High School: Pertevniyal Lycee (1964)
BS: Istanbul University. Science Licence Diploma in Zoology and Botany (1964-1968)
Graduate Study: The University of Kansas, Department of Entomology (1968-1969)
Ph.D.: State University of New York at Stony Brook. Ph.D. in Biological Sciences (1969-1973)
Advisor: Robert R. Sokal
Dissertation title: The effects of variation in larval development on laboratory populations of Tribolium and Houseflies.183 pp.
